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Scientific Papers

Peer-reviewed Journal Articles


Branch, T.A., I.M. Cȏté, S.R. David, J.A. Drew, M. LaRue, M.C. Márquez, E.C.M. Parsons, D. Rabaiotti, D. Shiffman, D.A. Steen, and A.L. Wild. 2024. Controlled experiment finds no detectable citation bump from Twitter promotion. PLOS ONE. In press.

Bogan, J.E., Jr., B.M. O’Hanlon, D.A. Steen, T. Horan, R. Taylor, A.K. Mason, T. Breen, H. Andreotta, B. Cornelius, A. Childress, and M. Elmore. 2024. Health assessment of free-ranging Eastern Indigo Snakes (Drymarchon couperi) from hydrologic restoration construction sites in South Florida, USA. Journal of Wildlife Diseases 60:39-51.


Chandler, H.C., D.A. Steen, J. Blue, J. E. Bogan, M. R. Bolt, T. Brady, D. R. Breininger, J. Buening, M. Elliott, J. Godwin, C. Guyer, R. L. Hill, M. Hoffman, N. L. Hyslop, C. L. Jenkins, C. Lechowicz, M. Moore, R. A. Moulis, S. Piccolomini, R. Redmond, F.H. Snow, B. S. Stegenga, D. J. Stevenson, J. Stiles, S. Stiles, M. Wallace, J. Waters, M. Wines, and J. M. Bauder. 2023. Evaluating growth rates of captive, wild, and reintroduced populations of the imperiled eastern indigo snake (Drymarchon couperi). Herpetologica 79:220-230.

Snyder, S.D., W.B. Sutton, and D.A. Steen. 2023. Species distribution modeling reveals insights into the occurrence of a locally rare snake at the periphery of its geographic range. Herpetologica 79:98-107.

Carr, J.L., J. Terry, D. Ligon, K.M. Enge, C.J. Krueger, T.M. Thomas, G. Brashear, G.R. Johnston, M. Gordon, G. Zwicky, E. Pierson Hill, D.J. Stevenson, and D.A. Steen. 2023. Nesting emergences and other terrestrial movements of Macrochelys (Testudines: Chelydridae). Southeastern Naturalist 22 Special Issue 12:378-387.


Jenkins, A.J., J.C. Godwin, D.A. Warner, and D.A. Steen. 2022. Movement ecology of Flattened Musk Turtles (Sternotherus depressus). Journal of Herpetology 56:1-7.


Goetz, S.M., D.A. Steen, M.A. Miller, C. Guyer, J. Kottwitz, J.F. Roberts, E. Blankenship, P.R. Pearson, D.A. Warner, and R.N. Reed. 2021. Argentine Black and White Tegu (Salvator merianae) can survive the winter under semi-natural conditions well beyond their current invasive range. PLoS One 16: e0245877.

Shamblin, B.M., M.G. Dodd, S.M. Pate, M.H. Godfrey, J.B. Pfaller, K.L. Williams, B.L. Ondich, D.A. Steen, E.S. Darrow, P. Hillbrand, R. Boettcher, M.S. Coyne, and C.J. Nairn. 2021. In search of the “missing majority” of nesting loggerhead turtles: improved inter-seasonal recapture rates through subpopulation-scale genetic tagging. Marine Biology 168 (16).


Barrett, K., C. Guyer, and D.A. Steen. 2020. The Evolutionary Community Concept is fully armed and operational: a reply to Sagoff. Biology and Philosophy 35:1-9.

Folt, B., C.P. McGowan, D.A. Steen, S. Piccolomini, M. Hoffman, J.C. Godwin, and C. Guyer. 2020. Modeling strategies and evaluating success during repatriations of elusive and endangered species. Animal Conservation 23:273-285.


Steen, D.A., S. Goetz, B. Ondich, M. Kaylor, A. Alterman, H. Weinstein, J. Gamby, J. Gorges, D.J. Stevenson, and T.M. Norton. 2019. An investigation of prey discrimination by neonate red-bellied mudsnakes (Farancia abacura). Florida Scientist 82:129-133.

DeGregorio, B.A., J.H. Sperry, D.G. Kovar, and D.A. Steen. 2019. Southern Flying Squirrels (Glaucomys volans) as major predators of birds in nest boxes in Conecuh National Forest, Alabama. Southeastern Naturalist 18:476-488.

Folt, B., J Bauder, S. Spear, D. Stevenson, M. Hoffman, J. Oaks, P.L. Wood Jr, C. Jenkins, D.A. Steen, and C. Guyer. 2019. Taxonomic and conservation implications of population genetic admixture, mito-nuclear discordance, and male-biased dispersal of a large endangered snake, Drymarchon couperi. PLoS ONE 14(3): e0214439.

Lambert, M., and D. A. Steen. 2019. (Re)examining the effects of urbanization on the population structure of freshwater turtles across the United States. Conservation Biology 33:1193-1196

Goetz, S.M., S. Piccolomini, M. Hoffman, J. Bogan, M. Holding, M.T. Mendonca, and D.A. Steen. 2019. Serum-based inhibition of pitviper venom by Eastern Indigo Snakes (Drymarchon couperi). Open Biology 8.


Graham, S.G., R. Kline, D. A. Steen, and C. Kelehear. 2018. Description of an extant salamander from the Gulf Coastal Plain of North America: the reticulated siren (Siren reticulata). PLOS ONE 13:e0207460.

Goetz, S. G., J. C. Godwin, M. Hoffman, F. Antonio, and D. A. Steen. 2018. Eastern Indigo Snakes exhibit a strong innate response to pit viper scent and an ontogenetic shift in their response to mouse scent. Herpetologica 74:152-158.


Steen, D. A., K. Barrett, E. Clarke, and C. Guyer. 2017. Conceptualizing communities as natural entities: a philosophical argument with basic and applied implications. Biology and Philosophy 32:1019-1034.

Steen, D. A., and O. Robinson. 2017. Estimating freshwater turtle mortality rates and population declines following hook ingestion. Conservation Biology 31:1333-1339.

Steen, D. A., and D. Kelly. 2017. Interspecific combat observed among viperid snakes. Ecology 98:1727-1728.

J. U. Van Dyke, C. M. B. Jachowski, D. A. Steen, B. P. Jackson, and W. A. Hopkins. 2017. Spatial differences in trace element bioaccumulation in turtles exposed to a partially remediated coal fly-ash spill. Environmental Toxicology and Chemistry 36:201-211.


Godwin, J. C., D. A. Steen, D. Werneke, and J. W. Armbruster. 2016. Two significant records of exotic neotropical freshwater fish observed in southern Alabama. Southeastern Naturalist 15:57-60.

Reed, R. N., M. W. Hopken, D. A. Steen, B. G. Falk, and A. J. Piaggio. 2016. Integrating early detection with DNA barcoding: species identification of a non-native monitor lizard (Squamata: Varanidae) carcass in Mississippi, USA. Management of Biological Invasions 7:193-197.

Steen, D. A., J. A. Stiles, S. H. Stiles, J. C. Godwin, and C. Guyer. 2016. Observations of feeding behavior by reintroduced Indigo Snakes in southern Alabama. Herpetological Review 47:11-13.


Steen, D. A., M. Barbour, C. J. W. McClure, K. P. Wray, J. N. Macey, and D. J. Stevenson. 2015. Landscape scale habitat selection of Harlequin Coralsnakes (Micrurus fulvius) in three large, protected areas in the southeastern United States. Copeia 2015:1037-1042.

Graham, S. P., D. A. Steen, M. A. Bailey, J. C. Godwin, J. Stiles, S. Stiles, T. Langkilde, and C. Guyer. 2015. The ampconecuhhibians and reptiles of Conecuh National Forest, Escambia and Covington Counties, Alabama. Alabama Museum of Natural History Bulletin 32:1-111.

Steen, D. A., and K. Barrett. 2015. Should states value species at the edge of their geographic range? Journal of Wildlife Management 79:872-876.

Steen, D. A., P. A. Osborne, M. Dovčiak, D. A. Patrick, and J. P. Gibbs. 2015. A preliminary investigation in the short-term effects of a prescribed fire on habitat quality for a snake assemblage. Herpetological Conservation and Biology 10:263-272

Steen, D. A., Van Dyke, J. U., B. P. Jackson, and W. A. Hopkins. 2015. Reproduction and hatchling performance in freshwater turtles associated with a remediated coal fly-ash spill. Environmental Research  138:38-48.

Jachowski, D. S., D. C. Kesler, D. A. Steen, and J. R. Walters. 2015. Redefining baselines in endangered species recovery. Journal of Wildlife Management 79:3-9.


Van Dyke, J. U., D. A. Steen, B. P. Jackson, and W. A. Hopkins. 2014. Maternal transfer and embryonic assimilation of trace elements in freshwater turtles after remediation of a coal fly-ash spill.Environmental Pollution 194:38-49.

Steen, D. A., B. C. Hopkins, J. U. Van Dyke, and W. A. Hopkins. 2014. Prevalence of ingested fish hooks in freshwater turtles from five rivers in the southeastern United States. PLoS ONE 9:e91368. doi:10.1371/journal.pone.0091368.

Steen, D. A., J. C. Godwin, C. J. W. McClure and M. Barbour. 2014. Informing management of endemic habitat specialists: multiscale habitat selection of the Red Hills Salamander. Journal of Wildlife Management 78:463-470.

Steen, D. A., C. J. W. McClure, W. B. Sutton, D. C. Rudolph, J. B. Pierce, J. R. Lee, L. L. Smith, B. B. Gregory, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2014. Copperheads are common where kingsnakes are not: relationships between the abundances of a predator and one of their prey. Herpetologica 70:69-76.

Sutton, W. B., Y. Wang, C. J. Schweitzer, and D. A. Steen. 2014. Lizard microhabitat and microclimate relationships in southeastern pine-hardwood forests managed with prescribed burning and thinning. Forest Science 60:180-190.

Steen, D. A., C. J. W. McClure, J. C. Brock, D. C. Rudolph, J. B. Pierce, J. R. Lee, W. J. Humphries, B. B. Gregory, W. B. Sutton, L. L. Smith, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2014. Snake co-occurrence patterns are best explained by habitat and hypothesized effects of interspecific interactions. Journal of Animal Ecology 83:286-295.



Steen, D. A., and D. S. Jachowski. 2013. Expanding shifting baseline syndrome to accommodate increasing abundances. Restoration Ecology 5:527-529. Cover photo.

Steen, D. A., D. J. Stevenson, J. C. Beane, J. D. Willson, M. J. Aresco, J. C. Godwin, S. P. Graham, L. L. Smith, J. M. Howze, D. C. Rudolph, J. B. Pierce, J. R. Lee, B. B. Gregory, J. Jensen, S. H. Stiles, J. A. Stiles, N. H. Nazdrowicz, and C. Guyer. 2013. Terrestrial movements of the red-bellied mudsnake (Farancia abacura) and rainbow snake (F. erytrogramma). Herpetological Review 44:208-213.

Steen, D. A., C. J. W. McClure, and S. P. Graham. 2013. Relative influence of weather and season on anuran calling activity. Canadian Journal of Zoology 91:462-467.


Steen, D. A., L. L. Smith, G. Morris, L. M. Conner, A. R. Litt, S. Pokswinski, and C. Guyer. 2013. Response of Aspidoscelis sexlineata (six-lined racerunner) to habitat restoration in fire-suppressed Pinus palustris (longleaf pine) sandhills. Restoration Ecology 21:457-463. Cover photo.

Steen, D. A., L. M. Conner, L. L. Smith, L. Provencher, J. K. Hiers, S. Pokswinski, B. Helms, and C. Guyer. 2013. Bird assemblage response to restoration of fire-suppressed longleaf pine sandhills. Ecological Applications 23:134-147.

Steen, D. A., L. L. Smith, L. M. Conner, A. R. Litt, L. Provencher, J. K. Hiers, S. Pokswinski, and C. Guyer. 2013. Reptile assemblage response to restoration of fire-suppressed longleaf pine sandhills. Ecological Applications 23:148-158.

Steen, D. A., C. J. W. McClure, L. L. Smith, B. J. Halstead, C. K. Dodd, Jr., W. B. Sutton, J. R. Lee, D. L. Baxley, W. J. Humphries, and C. Guyer. 2013. The effect of coachwhip presence on body size of North American racers suggests competition between these sympatric snakes. Journal of Zoology 289:86-93.

Smith, L. L., D. A. Steen, L. M. Conner, and J. C. Rutledge. 2013. Effects of predator exclusion on nest and hatchling survival in the gopher tortoise. Journal of Wildlife Management 77:352-358.



Steen, D. A., C. J. W. McClure, J. C. Brock, D. C. Rudolph, J. B. Pierce, J. R. Lee, W. J. Humphries, B. B. Gregory, W. B. Sutton, L. L.Smith, D. L. Baxley, D. J. Stevenson, and C. Guyer. 2012. Landscape-level influences of terrestrial snake occupancy within the southeastern United States. Ecological Applications 22:1084-1097.
          Highlighted in the Bulletin of The Ecological Society of America (2012) 93:352-353, with cover photo

Steen D. A., J. P. Gibbs, K. A. Buhlmann, J. L. Carr, B. W. Compton, J. D. Congdon, J. S. Doody, J. C. Godwin, K. L. Holcomb, D. R. Jackson, F. J. Janzen, G. Johnson, M. T. Jones, J.T. Lamer, T. A. Langen, M. V. Plummer, J. W. Rowe, R. A. Saumure, J. K. Tucker, and D. S. Wilson. 2012. Terrestrial habitat requirements of nesting freshwater turtles. Biological Conservation 150:121-128.

Graham, S. P., D. A. Steen, R. D. Birkhead, and C. Guyer. 2012. The amphibians and reptiles of Tuskegee National Forest, fig_4Macon County, Alabama. Alabama Museum of Natural History Bulletin 29:1-59.

Steen, D. A., M. Baragona, C. J. W. McClure, K. C. Jones, and L. L. Smith. 2012. Demography of a small population of loggerhead musk turtles (Sternotherus minor) in Okaloosa County, Florida. Florida Field Naturalist 40:47-55.

Graham, S. P., E. K. Timpe, A. M. Durso, D. A. Steen, W. B. Sutton, K. T. Nelson, G. J. Brown III, M. A. Connell, K. M. Gray, and J. C. Godwin. 2012. The second known contact zone between Plethodon websteri and P. ventralis, and additional county records for Bibb County, Alabama. Herpetological Review 43:311-312.


Steen, D. A., K. C. Jones, J. B. Jensen, B. K. Rincon, J. C. Godwin, and S. P. Graham. 2011. New amphibian and reptile county records for the Florida panhandle. Herpetological Review 42:576-577.


Steen, D. A., S. C. Sterrett, A. M. Heupel, and L. L. Smith. 2010. Snapping turtle, Chelydra serpentina, overland movements near the southeastern extent of its range. Georgia Journal of Science 68:196-201.

Steen, D. A., A. E. Rall McGee, S. M. Hermann, J. A. Stiles, S. H. Stiles, and C. Guyer. 2010. Effects of forest management on amphibians and reptiles: generalist species obscure trends among native forest associates. Open Environmental Sciences 4:24-30.

Steen, D. A., L. L. Smith, and M. A. Bailey. 2010. Suggested modifications to terrestrial box traps for snakes. Herpetological Review 41:320-321.

Graham, S. P., A. M. Durso, D. A. Steen, V. M. Johnson, S. K. Hoss, D. J. McMoran, K. T. Nelson, S. C. Cox, E. C. Susko, J. W. Shapiro, and J. R. McMillan. 2010. An additional bioblitz competition in northwest Georgia yields new county records. Herpetological Review 41:383-384.

Steen, D. A. 2010. Snakes in the grass: secretive natural histories defy both conventional and progressive statistics. Herpetological Conservation and Biology 5:183-188.

Linehan, J. M., L. L. Smith, and D. A. Steen. 2010. Ecology of the eastern kingsnake (Lampropeltis getula getula) in a longleaf pine (Pinus palustris) forest in southwestern Georgia. Herpetological Conservation and Biology 5:94-101.

Steen, D. A., J. M. Linehan, and L. L. Smith. 2010. Multiscale habitat selection and refuge use of common kingsnakes, Lampropeltis getula, in southwestern Georgia. Copeia 2010:227-231.

Steen, D. A., A. D. Steen, S. Pokswinski, S. P. Graham, and L. L. Smith. 2010. Snakes using stumpholes and windfall tree-associated subterranean structures in longleaf pine forests. Reptiles and Amphibians 17:49-51.

Graham, S. P., D. A. Steen, K. T. Nelson, A. M. Durso, and J. C. Maerz. 2010.  An overlooked hotspot? Rapid biodiversity assessment reveals a region of exceptional herpetofaunal richness in the southeastern United States.  Southeastern Naturalist 9:19-34.


Steen, D. A. and L. L. Smith. 2009. Eastern kingsnake, Lampropeltis getula getula, home ranges exhibit limited overlap. Southeastern Naturalist 8:553-558.

Graham, S. P., S. K. Hoss, R. D. Birkhead, C. K. Ward, D. A. Steen, K. M. Gray, and R. H. Mount. 2009. 101 new county records for Alabama herpetofauna. Herpetological Review 40:367-371.


Steen, D. A., L. L. Smith, L. M. Conner, J. C. Brock, and S. K. Hoss. 2007. Habitat use of sympatric rattlesnake species within the Gulf Coastal Plain. Journal of Wildlife Management 71:759-764.

Steen, D. A., S. C. Sterrett, S. A. Miller, and L. L. Smith. 2007. Terrestrial movements and microhabitat selection of overwintering subadult eastern mud turtles (Kinosternon subrubrum) in southwest Georgia. Journal of Herpetology 41:532-535.


Smith, L. L., D. A. Steen, J. M. Stober, M. C. Freeman, S. W. Golladay, L. M. Conner, and J. C. Cochrane. 2006. The vertebrate fauna of Ichauway, Baker County, Georgia. Southeastern Naturalist 5:599-620.

Steen, D. A., J. P. Gibbs and S. T. A. Timmermans. 2006. Assessing the sensitivity of waterbird communities to hydrologic change in the eastern Great Lakes region. Wetlands 26:605-611.

Steen, D. A. and L. L. Smith. 2006. Road surveys for turtles: consideration of possible sampling biases. Herpetological Conservation and Biology 1:9-15.

Steen, D. A., L. L. Smith, G. J. Miller, and S. C. Sterrett. 2006. Post breeding terrestrial movements of eastern tiger salamanders, Ambystoma tigrinum. Southeastern Naturalist 5:285-288.

Steen, D. A., M. J. Aresco, S. G. Beilke, B. W. Compton, E. P. Condon, C. K Dodd, Jr., H. Forrester, J. W. Gibbons, J. L. Greene, G. Johnson, T. A. Langen, M. J. Oldham, D. N. Oxier, R. A. Saumure, F. W. Schueler, J. M. Sleeman, L. L. Smith, J. K. Tucker and J. P. Gibbs. 2006. Relative vulnerability of female turtles to road mortality. Animal Conservation 9:269-273.


Gibbs, J. P. and D. A. Steen. 2005. Trends in sex ratios of turtles in the United States: implications of road mortality. Conservation Biology 19:552-556

Steen, D. A. and J. P. Gibbs. 2005. Potential influence of the terrestrial landscape on the distribution of pond-breeding eastern newts, Notophthalmus viridescens, within wetlands. Applied Herpetology 2:425-428.


Steen, D.A. and J. P. Gibbs. 2004. Effects of roads on the structure of freshwater turtle populations. Conservation Biology 18:1143-1148.


Jachowski, D. S., S. Bremner-Harrison, D. A. Steen, and K. Aarestrup. 2016. Accounting for potential physiological, behavioral, and community-level responses to reintroduction. In Reintroduction of Fish and Wildlife Populations (D. S. Jachowski, J. J. Millspaugh, P. L. Angermeier, and R. Slotow, eds.), University of California Press, Berkeley, CA, USA.

Gitzen, R., A., B. J. Keller, M. A. Miller, S. M. Goetz, D. A. Steen, D. S. Jachowski, J. C. Godwin, and J. J. Millspaugh. 2016. Effective and purposeful monitoring of species reintroductions. In Reintroduction of Fish and Wildlife Populations (D. S. Jachowski, J. J. Millspaugh, P. L. Angermeier, and R. Slotow, eds.), University of California Press, Berkeley, CA, USA.

Steen, D. A., C. Guyer, and L. L. Smith. 2012. A case study of relative abundance in snakes. Pages 287-294 In Reptile Biodiversity: Standard Methods for Inventory and Monitoring (R. W. McDiarmid, M. S. Foster, C. Guyer, J. W. Gibbons, and N. Chernoff, eds.), University of California Press, Berkeley, CA, USA.

Peer-reviewed Notes and Reviews

David, L.R., A.C. Hipps, K.M. Enge, and D.A. Steen. 2023. Nerodia fasciata (Southern Watersnake). Diet. Herpetological Review 54:316-317.

Hipps, A.C., K.M. Enge, and D.A. Steen. 2023. Pantherophis guttatus (Red Cornsnake). Attempted Predation. Herpetological Review 54:318.

Bogan Jr., J.E., D.A. Steen, B. O’Hanlon, M.M. Garner, H.D.S. Walden, and J.F.X. Wellehan, Jr. 2022. Drymarchon couperi (Eastern Indigo Snake). Death associated with Raillietiella orientalis. Herpetological Review 53:147.

Steen, D.A. 2021. Review of The Timber Rattlesnake: Life History, Distribution, Status, and Conservation Action Plan. Herpetological Review 52.

Broussard, M.A., J. Roach, M. Hoffman, and D.A. Steen. 2021. Drymarchon melanurus (Central American Indigo Snake). Behavior. Herpetological Review 52:154-155.

Broussard, M.A., J. Byers, and D.A. Steen. Incilius nebulifer (Gulf Coast Toad). Diet. Herpetological Review 52:69.

Greene, A.H., J. Levy, H. Levy, and D.A. Steen. 2020. Siren lacertina (Greater Siren). Geographic Distribution. Herpetological Review 51:75-76.

Enge, K.M., and D.A. Steen. 2020. Macrochelys suwanniensis (Suwannee Alligator Snapping Turtle). Geographic Distribution. Herpetological Review 51:772.

Payne, R., J.D. Mays, and D.A. Steen. 2020. Farancia erytrogramma (Rainbow Snake). Geographic Distribution. Herpetological Review 51:545.

Greene, A.H., D.B. Miller, D. Rubadou, R.H. Robins, and D.A. Steen. 2020. Nerodia clarkii compressicauda (Mangrove Saltmarsh Watersnake). Diet. Herpetological Review 51:871.

Broussard, M.A., A. Hetrick, C. Hetrick, B. Ondich, and D.A. Steen. 2020. Sonora semiannulata (Western Groundsnake). Behavior/Male-Male Combat. Herpetological Review 51:359-360.

Steen, D.A. 2019. Reviews of Dreaming in Turtle and Ecology and Conservation of the Diamond-Backed Terrapin. Biological Conservation 238:108194.

Watts, M.W, R. J. Emerson, J.E. Colbert, and D.A. Steen. 2019. Ophisaurus ventralis (Eastern Glass Lizard). Diet. Herpetological Review 50:788.

Lopel, L., D. J. Stevenson, and D. A. Steen. 2019. Plestiodon laticeps. (Broad-Headed Skink). Diet. Herpetological Review 50:789.

Paden, L., B.L. Ondich, T.M. Norton, and D. A. Steen. 2019. Pantherophis guttatus (Red Cornsnake). Attempted predation and defensive behavior. Hepetological Review 50:397.

Jarboe, C.A., J. E. Colbert, B. Zulkiewicz, and D. A. Steen. 2019. Osteopilus septentrionalis (Cuban Treefrog). Geographic Distribution. Herpetological Review 50:97-98.

Jarboe, C.A., J. E. Colbert, Y. Moore, D.A. Steen, and S. A. Johnson. 2019. Osteopilus septentrionalis (Cuban Treefrog). Geographic Distribution. Herpetological Review 50:98.

Butler, Z., B. Ondich, J. Gaskin, T. Norton, D. A. Steen, B. Sorg, and G. Bishop. 2019. Cemophora coccinea (Scarletsnake). Predatory Behavior. Herpetological Review 50:151.

Ondich, B.L., K. A. Mascovich, Z. Butler, and D. A. Steen. 2018. Pseudemys nelsoni (Florida Red-Bellied Cooter). Geographic Distribution. Herpetological Review 49:502.

Paden, L., B. L. Ondich, T. M. Norton, A. Alterman, and D. A. Steen. 2018. Pseudemys floridana (Florida Cooter). Geographic Distribution. Herpetological Review 49:712-713.

Paden, L., R. G. Bauer, B. L. Ondich, T. M. Norton, A. Alterman, and D. A. Steen. 2018. Pseudemys concinna (River Cooter). Geographic Distribution. Herpetological Review 49:712.

Hollister R., J. Hollister, S.R. David, and D. A. Steen. 2018. Crotalus oreganus (Western Rattlesnake). Predation. Herpetological Review 49:751.

Turk, E., A. Sloan, and D. A. Steen. 2018. Nerodia taxispilota (Brown Watersnake). Predation. Herpetological Review: 49:136.

Steen, D. A., J. U. Van Dyke, M. C. Dziadzio, T. D. Parker, H. J. Vogel, K. Q. Webb, and W. A. Hopkins. Kinosternon subrubrum (Eastern Mud Turtle). Geographic Distribution. Herpetological Review: in press.

Karlström, Å, and D. A. Steen. 2016. Nerodia cyclopion (Mississippi Green Watersnake). Geographic Distribution. Herpetological Review 47:631.

Grove, B., and D. A. Steen. 2015. Pantherophis guttatus (Red Cornsnake). Geographic Distribution. Herpetological Review 46:575.

Arbaugh, R, T. Arbaugh, and D. A. Steen. 2015. Nerodia erythrogaster (Plain-bellied Watersnake). Predation. Herpetological Review 46:647-648.

Hammerstein, G., D. A. Steen, and D. J. Stevenson. 2015. Crotalus horridus (Timber Rattlesnake). Diet. Herpetological Review 46:640-641.

Barrett, K., D. A. Steen, S. C. Sterrett, W. B. Sutton, and S. P. Graham. 2015. Desmognathus quadramaculatus (Black-bellied Salamander). Maximum Clutch Size. Herpetological Review 46:409.

Neal, T., and D. A. Steen. 2015. Agkistrodon piscivorus (Cottonmouth). Predation. Herpetological Review 46:264.

Graham, S. P., D. A. Steen, and D. Printiss. 2015. Gopherus polyphemus (Gopher Tortoise). Burrow Associate. Herpetological Review 46:244.

Steen, D. A. 2013. Pantherophis alleghaniensis (Eastern Ratsnake). Reproduction. Catesbeiana 33:69-70.

Steen, D. A. 2013. Clemmys guttata (Spotted Turtle). Reproduction. Herpetological Review 44:501.

Steen, D. A., M. C. Dziadzio, T. D. Parker, H. J. Vogel, K. Q. Webb, and W. A. Hopkins. 2012. Apalone mutica (Smooth Softshell). Geographic Distribution. Herpetological Review 43:614.

Steen, D. A., K. Q. Webb, H. J. Vogel, T. D. Parker, M. C. Dziadzio, and W. A. Hopkins. 2012. Kinosternon subrubrum (Eastern Mud Turtle). Diet. Herpetological Review 43:639.

Stevenson, D. J., D. A. Steen, and M. Wallace. 2012. Nerodia erythrogaster erythrogaster (Red-bellied Watersnake). Necrophagy by dung beetles (Scarabaeinae). Herpetological Review 43:347.

Steen, D. A., J. A. Stiles, S. H. Stiles, C. Guyer, J. B. Pierce, D. C. Rudolph, and L. L. Smith. 2011. Regina rigida (Glossy Crayfish Snake). Terrestrial Movement. Herpetological Review 42:102.

McClure, C. J. W., and D. A. Steen. 2010. Hemidactylus turcicus (Mediterranean Gecko). Geographic Distribution. Herpetological Review 41:512.

Jones, K. C, B. K. Rincon, and D. A. Steen. 2010. Bufo terrestris (Southern Toad). Predation. Herpetological Review 41:334-335.

Steen, D. A., G. G. Sorrell, N. J. Paris, K. J. Paris, D. D. Simpson, and L. L. Smith. 2010. Heterodon platirhinos (Eastern Hog-nosed Snake). Predator/prey mass ratio. Herpetological Review 41:365.

Heupel, A. M., K. E. McKean, J. M. Linehan, D. A. Steen, and L. L. Smith. 2009. Ambystoma sp. Habitat Use. Herpetological Review 40:66.

Tyson, A., D. A. Steen, L. L. Smith. 2008. Nerodia taxispilota (Brown Watersnake). Diet. Herpetological Review 39:472.

Muenz, T. K., D. A. Steen, B. E. Bass, and A. Kaeser. 2008. Apalone spinifera aspera (Gulf Coast Spiny Softshell). Predation. Herpetological Review 39:212.

Steen, D. A., G. J. Miller, S. C. Sterrett, and L. L. Smith. 2007. Masticophis flagellum (Eastern Coachwhip). Growth and Movement. Herpetological Review 38:90.

Sterrett, S. C., D. A. Steen, and L. L. Smith. 2006. Ambystoma tigrinum tigrinum  (Eastern Tiger Salamander). Burrow Description. Herpetological Review 37:435-436.

Steen, D. A., G. J. Miller, S. C. Sterrett, and L. L. Smith. 2005. Gopherus polyphemus (Gopher Tortoise). Juvenile use of gopher tunnel. Herpetological Review 46:442.

Steen, D. A., S. N. Becker, and L. L. Smith. 2005. Heterodon platirhinos (Eastern Hog-nosed Snake). Reproduction. Herpetological Review 46:457.

Steen, D.A., and J.P. Gibbs. 2004. Of roads and turtles: a summary of recent research findings. Turtle and Tortoise Newsletter 8:8-9.

White-faced Capuchin (Cebus capucinus) contemplating life and lunch in Palo Verde National Park, Costa Rica.